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lectures on evolution- µÚ4²¿·Ö


°´¼üÅÌÉÏ·½Ïò¼ü ¡û »ò ¡ú ¿É¿ìËÙÉÏÏ·­Ò³£¬°´¼üÅÌÉ쵀 Enter ¼ü¿É»Øµ½±¾ÊéĿ¼ҳ£¬°´¼üÅÌÉÏ·½Ïò¼ü ¡ü ¿É»Øµ½±¾Ò³¶¥²¿£¡

the¡¡huge¡¡mass¡¡of¡¡rocks¡¡extending¡¡from¡¡the¡¡middle¡¡of¡¡the

Palaeozoic¡¡formations¡¡to¡¡the¡¡uppermost¡¡members¡¡of¡¡the¡¡series£»

must¡¡belong¡¡to¡¡the¡¡day¡¡which¡¡is¡¡termed¡¡by¡¡Milton¡¡the¡¡sixth¡£

But£»¡¡further£»¡¡it¡¡is¡¡expressly¡¡stated¡¡that¡¡aquatic¡¡animals¡¡took

their¡¡origin¡¡on¡¡the¡¡fifth¡¡day£»¡¡and¡¡not¡¡before£»¡¡hence£»¡¡all

formations¡¡in¡¡which¡¡remains¡¡of¡¡aquatic¡¡animals¡¡can¡¡be¡¡proved¡¡to

exist£»¡¡and¡¡which¡¡therefore¡¡testify¡¡that¡¡such¡¡animals¡¡lived¡¡at

the¡¡time¡¡when¡¡these¡¡formations¡¡were¡¡in¡¡course¡¡of¡¡deposition£»

must¡¡have¡¡been¡¡deposited¡¡during¡¡or¡¡since¡¡the¡¡period¡¡which¡¡Milton

speaks¡¡of¡¡as¡¡the¡¡fifth¡¡day¡£¡¡But¡¡there¡¡is¡¡absolutely¡¡no

fossiliferous¡¡formation¡¡in¡¡which¡¡the¡¡remains¡¡of¡¡aquatic¡¡animals

are¡¡absent¡£¡¡The¡¡oldest¡¡fossils¡¡in¡¡the¡¡Silurian¡¡rocks¡¡are¡¡exuviae

of¡¡marine¡¡animals£»¡¡and¡¡if¡¡the¡¡view¡¡which¡¡is¡¡entertained¡¡by

Principal¡¡Dawson¡¡and¡¡Dr¡£¡¡Carpenter¡¡respecting¡¡the¡¡nature¡¡of¡¡the

Eozoon¡¡be¡¡well¡­founded£»¡¡aquatic¡¡animals¡¡existed¡¡at¡¡a

period¡¡as¡¡far¡¡antecedent¡¡to¡¡the¡¡deposition¡¡of¡¡the¡¡coal¡¡as¡¡the

coal¡¡is¡¡from¡¡us£»¡¡inasmuch¡¡as¡¡the¡¡Eozoon¡¡is¡¡met¡¡with¡¡in

those¡¡Laurentian¡¡strata¡¡which¡¡lie¡¡at¡¡the¡¡bottom¡¡of¡¡the¡¡series¡¡of

stratified¡¡rocks¡£¡¡Hence¡¡it¡¡follows£»¡¡plainly¡¡enough£»¡¡that¡¡the

whole¡¡series¡¡of¡¡stratified¡¡rocks£»¡¡if¡¡they¡¡are¡¡to¡¡be¡¡brought¡¡into

harmony¡¡with¡¡Milton£»¡¡must¡¡be¡¡referred¡¡to¡¡the¡¡fifth¡¡and¡¡sixth

days£»¡¡and¡¡that¡¡we¡¡cannot¡¡hope¡¡to¡¡find¡¡the¡¡slightest¡¡trace¡¡of¡¡the

products¡¡of¡¡the¡¡earlier¡¡days¡¡in¡¡the¡¡geological¡¡record¡£¡¡When¡¡we

consider¡¡these¡¡simple¡¡facts£»¡¡we¡¡see¡¡how¡¡absolutely¡¡futile¡¡are

the¡¡attempts¡¡that¡¡have¡¡been¡¡made¡¡to¡¡draw¡¡a¡¡parallel¡¡between¡¡the

story¡¡told¡¡by¡¡so¡¡much¡¡of¡¡the¡¡crust¡¡of¡¡the¡¡earth¡¡as¡¡is¡¡known¡¡to

us¡¡and¡¡the¡¡story¡¡which¡¡Milton¡¡tells¡£¡¡The¡¡whole¡¡series¡¡of

fossiliferous¡¡stratified¡¡rocks¡¡must¡¡be¡¡referred¡¡to¡¡the¡¡last¡¡two

days£»¡¡and¡¡neither¡¡the¡¡Carboniferous£»¡¡nor¡¡any¡¡other£»¡¡formation

can¡¡afford¡¡evidence¡¡of¡¡the¡¡work¡¡of¡¡the¡¡third¡¡day¡£



Not¡¡only¡¡is¡¡there¡¡this¡¡objection¡¡to¡¡any¡¡attempt¡¡to¡¡establish¡¡a

harmony¡¡between¡¡the¡¡Miltonic¡¡account¡¡and¡¡the¡¡facts¡¡recorded¡¡in

the¡¡fossiliferous¡¡rocks£»¡¡but¡¡there¡¡is¡¡a¡¡further¡¡difficulty¡£

According¡¡to¡¡the¡¡Miltonic¡¡account£»¡¡the¡¡order¡¡in¡¡which¡¡animals

should¡¡have¡¡made¡¡their¡¡appearance¡¡in¡¡the¡¡stratified¡¡rocks¡¡would

be¡¡thus£º¡¡Fishes£»¡¡including¡¡the¡¡great¡¡whales£»¡¡and¡¡birds£»

after¡¡them£»¡¡all¡¡varieties¡¡of¡¡terrestrial¡¡animals¡¡except¡¡birds¡£

Nothing¡¡could¡¡be¡¡further¡¡from¡¡the¡¡facts¡¡as¡¡we¡¡find¡¡them£»¡¡we¡¡know

of¡¡not¡¡the¡¡slightest¡¡evidence¡¡of¡¡the¡¡existence¡¡of¡¡birds¡¡before

the¡¡Jurassic£»¡¡or¡¡perhaps¡¡the¡¡Triassic£»¡¡formation£»

while¡¡terrestrial¡¡animals£»¡¡as¡¡we¡¡have¡¡just¡¡seen£»¡¡occur¡¡in¡¡the

Carboniferous¡¡rocks¡£



If¡¡there¡¡were¡¡any¡¡harmony¡¡between¡¡the¡¡Miltonic¡¡account¡¡and¡¡the

circumstantial¡¡evidence£»¡¡we¡¡ought¡¡to¡¡have¡¡abundant¡¡evidence¡¡of

the¡¡existence¡¡of¡¡birds¡¡in¡¡the¡¡Carboniferous£»¡¡the¡¡Devonian£»¡¡and

the¡¡Silurian¡¡rocks¡£¡¡I¡¡need¡¡hardly¡¡say¡¡that¡¡this¡¡is¡¡not¡¡the¡¡case£»

and¡¡that¡¡not¡¡a¡¡trace¡¡of¡¡birds¡¡makes¡¡its¡¡appearance¡¡until¡¡the¡¡far

later¡¡period¡¡which¡¡I¡¡have¡¡mentioned¡£



And¡¡again£»¡¡if¡¡it¡¡be¡¡true¡¡that¡¡all¡¡varieties¡¡of¡¡fishes¡¡and¡¡the

great¡¡whales£»¡¡and¡¡the¡¡like£»¡¡made¡¡their¡¡appearance¡¡on¡¡the¡¡fifth

day£»¡¡we¡¡ought¡¡to¡¡find¡¡the¡¡remains¡¡of¡¡these¡¡animals¡¡in¡¡the¡¡older

rocksin¡¡those¡¡which¡¡were¡¡deposited¡¡before¡¡the¡¡Carboniferous

epoch¡£¡¡Fishes¡¡we¡¡do¡¡find£»¡¡in¡¡considerable¡¡number¡¡and¡¡variety£»

but¡¡the¡¡great¡¡whales¡¡are¡¡absent£»¡¡and¡¡the¡¡fishes¡¡are¡¡not¡¡such¡¡as

now¡¡live¡£¡¡Not¡¡one¡¡solitary¡¡species¡¡of¡¡fish¡¡now¡¡in¡¡existence¡¡is

to¡¡be¡¡found¡¡in¡¡the¡¡Devonian¡¡or¡¡Silurian¡¡formations¡£¡¡Hence¡¡we¡¡are

introduced¡¡afresh¡¡to¡¡the¡¡dilemma¡¡which¡¡I¡¡have¡¡already¡¡placed

before¡¡you£º¡¡either¡¡the¡¡animals¡¡which¡¡came¡¡into¡¡existence¡¡on¡¡the

fifth¡¡day¡¡were¡¡not¡¡such¡¡as¡¡those¡¡which¡¡are¡¡found¡¡at¡¡present£»¡¡are

not¡¡the¡¡direct¡¡and¡¡immediate¡¡ancestors¡¡of¡¡those¡¡which¡¡now¡¡exist£»

in¡¡which¡¡case£»¡¡either¡¡fresh¡¡creations¡¡of¡¡which¡¡nothing¡¡is¡¡said£»

or¡¡a¡¡process¡¡of¡¡evolution£»¡¡must¡¡have¡¡occurred£»¡¡or¡¡else¡¡the¡¡whole

story¡¡must¡¡be¡¡given¡¡up£»¡¡as¡¡not¡¡only¡¡devoid¡¡of¡¡any¡¡circumstantial

evidence£»¡¡but¡¡contrary¡¡to¡¡such¡¡evidence¡¡as¡¡exists¡£



I¡¡placed¡¡before¡¡you¡¡in¡¡a¡¡few¡¡words£»¡¡some¡¡little¡¡time¡¡ago£»¡¡a

statement¡¡of¡¡the¡¡sum¡¡and¡¡substance¡¡of¡¡Milton's¡¡hypothesis¡£

Let¡¡me¡¡now¡¡try¡¡to¡¡state¡¡as¡¡briefly£»¡¡the¡¡effect¡¡of¡¡the

circumstantial¡¡evidence¡¡bearing¡¡upon¡¡the¡¡past¡¡history¡¡of¡¡the

earth¡¡which¡¡is¡¡furnished£»¡¡without¡¡the¡¡possibility¡¡of¡¡mistake£»

with¡¡no¡¡chance¡¡of¡¡error¡¡as¡¡to¡¡its¡¡chief¡¡features£»¡¡by¡¡the

stratified¡¡rocks¡£¡¡What¡¡we¡¡find¡¡is£»¡¡that¡¡the¡¡great¡¡series¡¡of

formations¡¡represents¡¡a¡¡period¡¡of¡¡time¡¡of¡¡which¡¡our¡¡human

chronologies¡¡hardly¡¡afford¡¡us¡¡a¡¡unit¡¡of¡¡measure¡£¡¡I¡¡will¡¡not

pretend¡¡to¡¡say¡¡how¡¡we¡¡ought¡¡to¡¡estimate¡¡this¡¡time£»¡¡in¡¡millions

or¡¡in¡¡billions¡¡of¡¡years¡£¡¡For¡¡my¡¡purpose£»¡¡the¡¡determination¡¡of

its¡¡absolute¡¡duration¡¡is¡¡wholly¡¡unessential¡£¡¡But¡¡that¡¡the¡¡time

was¡¡enormous¡¡there¡¡can¡¡be¡¡no¡¡question¡£



It¡¡results¡¡from¡¡the¡¡simplest¡¡methods¡¡of¡¡interpretation£»¡¡that

leaving¡¡out¡¡of¡¡view¡¡certain¡¡patches¡¡of¡¡metamorphosed¡¡rocks£»¡¡and

certain¡¡volcanic¡¡products£»¡¡all¡¡that¡¡is¡¡now¡¡dry¡¡land¡¡has¡¡once

been¡¡at¡¡the¡¡bottom¡¡of¡¡the¡¡waters¡£¡¡It¡¡is¡¡perfectly¡¡certain¡¡that£»

at¡¡a¡¡comparatively¡¡recent¡¡period¡¡of¡¡the¡¡world's¡¡historythe

Cretaceous¡¡epochnone¡¡of¡¡the¡¡great¡¡physical¡¡features¡¡which¡¡at

present¡¡mark¡¡the¡¡surface¡¡of¡¡the¡¡globe¡¡existed¡£¡¡It¡¡is¡¡certain

that¡¡the¡¡Rocky¡¡Mountains¡¡were¡¡not¡£¡¡It¡¡is¡¡certain¡¡that¡¡the

Himalaya¡¡Mountains¡¡were¡¡not¡£¡¡It¡¡is¡¡certain¡¡that¡¡the¡¡Alps¡¡and¡¡the

Pyrenees¡¡had¡¡no¡¡existence¡£¡¡The¡¡evidence¡¡is¡¡of¡¡the¡¡plainest

possible¡¡character¡¡and¡¡is¡¡simply¡¡this£ºWe¡¡find¡¡raised¡¡up¡¡on¡¡the

flanks¡¡of¡¡these¡¡mountains£»¡¡elevated¡¡by¡¡the¡¡forces¡¡of¡¡upheaval

which¡¡have¡¡given¡¡rise¡¡to¡¡them£»¡¡masses¡¡of¡¡Cretaceous¡¡rock¡¡which

formed¡¡the¡¡bottom¡¡of¡¡the¡¡sea¡¡before¡¡those¡¡mountains¡¡existed¡£

It¡¡is¡¡therefore¡¡clear¡¡that¡¡the¡¡elevatory¡¡forces¡¡which¡¡gave¡¡rise

to¡¡the¡¡mountains¡¡operated¡¡subsequently¡¡to¡¡the¡¡Cretaceous¡¡epoch£»

and¡¡that¡¡the¡¡mountains¡¡themselves¡¡are¡¡largely¡¡made¡¡up¡¡of¡¡the

materials¡¡deposited¡¡in¡¡the¡¡sea¡¡which¡¡once¡¡occupied¡¡their¡¡place¡£

As¡¡we¡¡go¡¡back¡¡in¡¡time£»¡¡we¡¡meet¡¡with¡¡constant¡¡alternations¡¡of¡¡sea

and¡¡land£»¡¡of¡¡estuary¡¡and¡¡open¡¡ocean£»¡¡and£»¡¡in¡¡correspondence¡¡with

these¡¡alternations£»¡¡we¡¡observe¡¡the¡¡changes¡¡in¡¡the¡¡fauna¡¡and

flora¡¡to¡¡which¡¡I¡¡have¡¡referred¡£



But¡¡the¡¡inspection¡¡of¡¡these¡¡changes¡¡gives¡¡us¡¡no¡¡right¡¡to¡¡believe

that¡¡there¡¡has¡¡been¡¡any¡¡discontinuity¡¡in¡¡natural¡¡processes¡£

There¡¡is¡¡no¡¡trace¡¡of¡¡general¡¡cataclysms£»¡¡of¡¡universal¡¡deluges£»

or¡¡sudden¡¡destructions¡¡of¡¡a¡¡whole¡¡fauna¡¡or¡¡flora¡£

The¡¡appearances¡¡which¡¡were¡¡formerly¡¡interpreted¡¡in¡¡that¡¡way¡¡have

all¡¡been¡¡shown¡¡to¡¡be¡¡delusive£»¡¡as¡¡our¡¡knowledge¡¡has¡¡increased

and¡¡as¡¡the¡¡blanks¡¡which¡¡formerly¡¡appeared¡¡to¡¡exist¡¡between¡¡the

different¡¡formations¡¡have¡¡been¡¡filled¡¡up¡£¡¡That¡¡there¡¡is¡¡no

absolute¡¡break¡¡between¡¡formation¡¡and¡¡formation£»¡¡that¡¡there¡¡has

been¡¡no¡¡sudden¡¡disappearance¡¡of¡¡all¡¡the¡¡forms¡¡of¡¡life¡¡and

replacement¡¡of¡¡them¡¡by¡¡others£»¡¡but¡¡that¡¡changes¡¡have¡¡gone¡¡on

slowly¡¡and¡¡gradually£»¡¡that¡¡one¡¡type¡¡has¡¡died¡¡out¡¡and¡¡another¡¡has

taken¡¡its¡¡place£»¡¡and¡¡that¡¡thus£»¡¡by¡¡insensible¡¡degrees£»¡¡one¡¡fauna

has¡¡been¡¡replaced¡¡by¡¡another£»¡¡are¡¡conclusions¡¡strengthened¡¡by

constantly¡¡increasing¡¡evidence¡£¡¡So¡¡that¡¡within¡¡the¡¡whole¡¡of¡¡the

immense¡¡period¡¡indicated¡¡by¡¡the¡¡fossiliferous¡¡stratified¡¡rocks£»

there¡¡is¡¡assuredly¡¡not¡¡the¡¡slightest¡¡proof¡¡of¡¡any¡¡break¡¡in¡¡the

uniformity¡¡of¡¡Nature's¡¡operations£»¡¡no¡¡indication¡¡that¡¡events

have¡¡followed¡¡other¡¡than¡¡a¡¡clear¡¡and¡¡orderly¡¡sequence¡£



That£»¡¡I¡¡say£»¡¡is¡¡the¡¡natural¡¡and¡¡obvious¡¡teaching¡¡of¡¡the

circumstantial¡¡evidence¡¡contained¡¡in¡¡the¡¡stratified¡¡rocks¡£¡¡I

leave¡¡you¡¡to¡¡consider¡¡how¡¡far£»¡¡by¡¡any¡¡ingenuity¡¡of

interpretation£»¡¡by¡¡any¡¡stretching¡¡of¡¡the¡¡meaning¡¡of¡¡language£»¡¡it

can¡¡be¡¡brought¡¡into¡¡harmony¡¡with¡¡the¡¡Miltonic¡¡hypothesis¡£



There¡¡remains¡¡the¡¡third¡¡hypothesis£»¡¡that¡¡of¡¡which¡¡I¡¡have¡¡spoken

as¡¡the¡¡hypothesis¡¡of¡¡evolution£»¡¡and¡¡I¡¡purpose¡¡that£»¡¡in¡¡lectures

to¡¡come£»¡¡we¡¡should¡¡discuss¡¡it¡¡as¡¡carefully¡¡as¡¡we¡¡have¡¡considered

the¡¡other¡¡two¡¡hypotheses¡£¡¡I¡¡need¡¡not¡¡say¡¡that¡¡it¡¡is¡¡quite

hopeless¡¡to¡¡look¡¡for¡¡testimonial¡¡evidence¡¡of¡¡evolution¡£¡¡The¡¡very

nature¡¡of¡¡the¡¡case¡¡precludes¡¡the¡¡possibility¡¡of¡¡such¡¡evidence£»

for¡¡the¡¡human¡¡race¡¡can¡¡no¡¡more¡¡be¡¡expected¡¡to¡¡testify¡¡to¡¡its¡¡own

origin£»¡¡than¡¡a¡¡child¡¡can¡¡be¡¡tendered¡¡as¡¡a¡¡witness¡¡of¡¡its¡¡own

birth¡£¡¡Our¡¡sole¡¡inquiry¡¡is£»¡¡what¡¡foundation¡¡circumstantial

evidence¡¡lends¡¡to¡¡the¡¡hypothesis£»¡¡or¡¡whether¡¡it¡¡lends¡¡none£»¡¡or

whether¡¡it¡¡controverts¡¡the¡¡hypothesis¡£¡¡I¡¡shall¡¡deal¡¡with¡¡the

matter¡¡entirely¡¡as¡¡a¡¡question¡¡of¡¡history¡£¡¡I¡¡shall¡¡not¡¡indulge¡¡in

the¡¡discussion¡¡of¡¡any¡¡speculative¡¡probabilities¡£¡¡I¡¡shall¡¡not

attempt¡¡to¡¡show¡¡that¡¡Nature¡¡is¡¡unintelligible¡¡unless¡¡we¡¡adopt

some¡¡such¡¡hypothesis¡£¡¡For¡¡anything¡¡I¡¡know¡¡about¡¡the¡¡matter£»¡¡it

may¡¡be¡¡the¡¡way¡¡of¡¡Nature¡¡to¡¡be¡¡unintelligible£»¡¡she¡¡is¡¡often

puzzling£»¡¡and¡¡I¡¡have¡¡no¡¡reason¡¡to¡¡suppose¡¡that¡¡she¡¡is¡¡bound¡¡to

fit¡¡herself¡¡to¡¡our¡¡notions¡£



I¡¡shall¡¡place¡¡before¡¡you¡¡three¡¡kinds¡¡of¡¡evidence¡¡entirely¡¡based

upon¡¡what¡¡is¡¡known¡¡of¡¡the¡¡forms¡¡of¡¡animal¡¡life¡¡which¡¡are

contained¡¡in¡¡the¡¡series¡¡of¡¡stratified¡¡rocks¡£¡¡I¡¡shall¡¡endeavour

to¡¡show¡¡you¡¡that¡¡there¡¡is¡¡one¡¡kind¡¡of¡¡evidence¡¡which¡¡is¡¡neutral£»

which¡¡neither¡¡helps¡¡evolution¡¡nor¡¡is¡¡inconsistent¡¡with¡¡it¡£

I¡¡shall¡¡then¡¡bring¡¡forward¡¡a¡¡second¡¡kind¡¡of¡¡evidence¡¡which

indicates¡¡a¡¡strong¡¡probability¡¡in¡¡favour¡¡of¡¡evolution£»¡¡but¡¡does

not¡¡prove¡¡it£»¡¡and£»¡¡lastly£»¡¡I¡¡shall¡¡adduce¡¡a¡¡third¡¡kind¡¡of

evidence¡¡which£»¡¡being¡¡as¡¡complete¡¡as¡¡any¡¡evidence¡¡which¡¡we¡¡can

hope¡¡to¡¡obtain¡¡upon¡¡such¡¡a¡¡subject£»¡¡and¡¡being¡¡wholly¡¡and

strikingly¡¡in¡¡favour¡¡of¡¡evolution£»¡¡may¡¡fairly¡¡be¡¡called

demonstrative¡¡evidence¡¡of¡¡its¡¡occurrence¡£







LECTURES¡¡ON¡¡EVOLUTION

II



THE¡¡HYPOTHESIS¡¡OF¡¡EVOLUTION¡£¡¡THE¡¡NEUTRAL¡¡AND

THE¡¡FAVOURABLE¡¡EVIDENCE¡£



In¡¡the¡¡preceding¡¡lecture¡¡I¡¡pointed¡¡out¡¡that¡¡there¡¡are¡¡three

hypotheses¡¡which¡¡may¡¡be¡¡entertained£»¡¡and¡¡which¡¡have¡¡been

entertained£»¡¡respecting¡¡the¡¡past¡¡history¡¡of¡¡life¡¡upon¡¡the¡¡globe¡£

According¡¡to¡¡the¡¡first¡¡of¡¡these¡¡hypotheses£»¡¡living¡¡beings£»¡¡such

as¡¡now¡¡exist£»¡¡have¡¡existed¡¡from¡¡all¡¡eternity¡¡upon¡¡this¡¡earth¡£

We¡¡tested¡¡that¡¡hypothesis¡¡by¡¡the¡¡circumstantial¡¡evidence£»¡¡as¡¡I

called¡¡it£»¡¡which¡¡is¡¡furnished¡¡by¡¡the¡¡fossil¡¡remains¡¡contained¡¡in

the¡¡earth's¡¡c
СÌáʾ£º°´ »Ø³µ [Enter] ¼ü ·µ»ØÊéÄ¿£¬°´ ¡û ¼ü ·µ»ØÉÏÒ»Ò³£¬ °´ ¡ú ¼ü ½øÈëÏÂÒ»Ò³¡£ ÔÞһϠÌí¼ÓÊéÇ©¼ÓÈëÊé¼Ü