the¡¡huge¡¡mass¡¡of¡¡rocks¡¡extending¡¡from¡¡the¡¡middle¡¡of¡¡the
Palaeozoic¡¡formations¡¡to¡¡the¡¡uppermost¡¡members¡¡of¡¡the¡¡series£»
must¡¡belong¡¡to¡¡the¡¡day¡¡which¡¡is¡¡termed¡¡by¡¡Milton¡¡the¡¡sixth¡£
But£»¡¡further£»¡¡it¡¡is¡¡expressly¡¡stated¡¡that¡¡aquatic¡¡animals¡¡took
their¡¡origin¡¡on¡¡the¡¡fifth¡¡day£»¡¡and¡¡not¡¡before£»¡¡hence£»¡¡all
formations¡¡in¡¡which¡¡remains¡¡of¡¡aquatic¡¡animals¡¡can¡¡be¡¡proved¡¡to
exist£»¡¡and¡¡which¡¡therefore¡¡testify¡¡that¡¡such¡¡animals¡¡lived¡¡at
the¡¡time¡¡when¡¡these¡¡formations¡¡were¡¡in¡¡course¡¡of¡¡deposition£»
must¡¡have¡¡been¡¡deposited¡¡during¡¡or¡¡since¡¡the¡¡period¡¡which¡¡Milton
speaks¡¡of¡¡as¡¡the¡¡fifth¡¡day¡£¡¡But¡¡there¡¡is¡¡absolutely¡¡no
fossiliferous¡¡formation¡¡in¡¡which¡¡the¡¡remains¡¡of¡¡aquatic¡¡animals
are¡¡absent¡£¡¡The¡¡oldest¡¡fossils¡¡in¡¡the¡¡Silurian¡¡rocks¡¡are¡¡exuviae
of¡¡marine¡¡animals£»¡¡and¡¡if¡¡the¡¡view¡¡which¡¡is¡¡entertained¡¡by
Principal¡¡Dawson¡¡and¡¡Dr¡£¡¡Carpenter¡¡respecting¡¡the¡¡nature¡¡of¡¡the
Eozoon¡¡be¡¡well¡founded£»¡¡aquatic¡¡animals¡¡existed¡¡at¡¡a
period¡¡as¡¡far¡¡antecedent¡¡to¡¡the¡¡deposition¡¡of¡¡the¡¡coal¡¡as¡¡the
coal¡¡is¡¡from¡¡us£»¡¡inasmuch¡¡as¡¡the¡¡Eozoon¡¡is¡¡met¡¡with¡¡in
those¡¡Laurentian¡¡strata¡¡which¡¡lie¡¡at¡¡the¡¡bottom¡¡of¡¡the¡¡series¡¡of
stratified¡¡rocks¡£¡¡Hence¡¡it¡¡follows£»¡¡plainly¡¡enough£»¡¡that¡¡the
whole¡¡series¡¡of¡¡stratified¡¡rocks£»¡¡if¡¡they¡¡are¡¡to¡¡be¡¡brought¡¡into
harmony¡¡with¡¡Milton£»¡¡must¡¡be¡¡referred¡¡to¡¡the¡¡fifth¡¡and¡¡sixth
days£»¡¡and¡¡that¡¡we¡¡cannot¡¡hope¡¡to¡¡find¡¡the¡¡slightest¡¡trace¡¡of¡¡the
products¡¡of¡¡the¡¡earlier¡¡days¡¡in¡¡the¡¡geological¡¡record¡£¡¡When¡¡we
consider¡¡these¡¡simple¡¡facts£»¡¡we¡¡see¡¡how¡¡absolutely¡¡futile¡¡are
the¡¡attempts¡¡that¡¡have¡¡been¡¡made¡¡to¡¡draw¡¡a¡¡parallel¡¡between¡¡the
story¡¡told¡¡by¡¡so¡¡much¡¡of¡¡the¡¡crust¡¡of¡¡the¡¡earth¡¡as¡¡is¡¡known¡¡to
us¡¡and¡¡the¡¡story¡¡which¡¡Milton¡¡tells¡£¡¡The¡¡whole¡¡series¡¡of
fossiliferous¡¡stratified¡¡rocks¡¡must¡¡be¡¡referred¡¡to¡¡the¡¡last¡¡two
days£»¡¡and¡¡neither¡¡the¡¡Carboniferous£»¡¡nor¡¡any¡¡other£»¡¡formation
can¡¡afford¡¡evidence¡¡of¡¡the¡¡work¡¡of¡¡the¡¡third¡¡day¡£
Not¡¡only¡¡is¡¡there¡¡this¡¡objection¡¡to¡¡any¡¡attempt¡¡to¡¡establish¡¡a
harmony¡¡between¡¡the¡¡Miltonic¡¡account¡¡and¡¡the¡¡facts¡¡recorded¡¡in
the¡¡fossiliferous¡¡rocks£»¡¡but¡¡there¡¡is¡¡a¡¡further¡¡difficulty¡£
According¡¡to¡¡the¡¡Miltonic¡¡account£»¡¡the¡¡order¡¡in¡¡which¡¡animals
should¡¡have¡¡made¡¡their¡¡appearance¡¡in¡¡the¡¡stratified¡¡rocks¡¡would
be¡¡thus£º¡¡Fishes£»¡¡including¡¡the¡¡great¡¡whales£»¡¡and¡¡birds£»
after¡¡them£»¡¡all¡¡varieties¡¡of¡¡terrestrial¡¡animals¡¡except¡¡birds¡£
Nothing¡¡could¡¡be¡¡further¡¡from¡¡the¡¡facts¡¡as¡¡we¡¡find¡¡them£»¡¡we¡¡know
of¡¡not¡¡the¡¡slightest¡¡evidence¡¡of¡¡the¡¡existence¡¡of¡¡birds¡¡before
the¡¡Jurassic£»¡¡or¡¡perhaps¡¡the¡¡Triassic£»¡¡formation£»
while¡¡terrestrial¡¡animals£»¡¡as¡¡we¡¡have¡¡just¡¡seen£»¡¡occur¡¡in¡¡the
Carboniferous¡¡rocks¡£
If¡¡there¡¡were¡¡any¡¡harmony¡¡between¡¡the¡¡Miltonic¡¡account¡¡and¡¡the
circumstantial¡¡evidence£»¡¡we¡¡ought¡¡to¡¡have¡¡abundant¡¡evidence¡¡of
the¡¡existence¡¡of¡¡birds¡¡in¡¡the¡¡Carboniferous£»¡¡the¡¡Devonian£»¡¡and
the¡¡Silurian¡¡rocks¡£¡¡I¡¡need¡¡hardly¡¡say¡¡that¡¡this¡¡is¡¡not¡¡the¡¡case£»
and¡¡that¡¡not¡¡a¡¡trace¡¡of¡¡birds¡¡makes¡¡its¡¡appearance¡¡until¡¡the¡¡far
later¡¡period¡¡which¡¡I¡¡have¡¡mentioned¡£
And¡¡again£»¡¡if¡¡it¡¡be¡¡true¡¡that¡¡all¡¡varieties¡¡of¡¡fishes¡¡and¡¡the
great¡¡whales£»¡¡and¡¡the¡¡like£»¡¡made¡¡their¡¡appearance¡¡on¡¡the¡¡fifth
day£»¡¡we¡¡ought¡¡to¡¡find¡¡the¡¡remains¡¡of¡¡these¡¡animals¡¡in¡¡the¡¡older
rocksin¡¡those¡¡which¡¡were¡¡deposited¡¡before¡¡the¡¡Carboniferous
epoch¡£¡¡Fishes¡¡we¡¡do¡¡find£»¡¡in¡¡considerable¡¡number¡¡and¡¡variety£»
but¡¡the¡¡great¡¡whales¡¡are¡¡absent£»¡¡and¡¡the¡¡fishes¡¡are¡¡not¡¡such¡¡as
now¡¡live¡£¡¡Not¡¡one¡¡solitary¡¡species¡¡of¡¡fish¡¡now¡¡in¡¡existence¡¡is
to¡¡be¡¡found¡¡in¡¡the¡¡Devonian¡¡or¡¡Silurian¡¡formations¡£¡¡Hence¡¡we¡¡are
introduced¡¡afresh¡¡to¡¡the¡¡dilemma¡¡which¡¡I¡¡have¡¡already¡¡placed
before¡¡you£º¡¡either¡¡the¡¡animals¡¡which¡¡came¡¡into¡¡existence¡¡on¡¡the
fifth¡¡day¡¡were¡¡not¡¡such¡¡as¡¡those¡¡which¡¡are¡¡found¡¡at¡¡present£»¡¡are
not¡¡the¡¡direct¡¡and¡¡immediate¡¡ancestors¡¡of¡¡those¡¡which¡¡now¡¡exist£»
in¡¡which¡¡case£»¡¡either¡¡fresh¡¡creations¡¡of¡¡which¡¡nothing¡¡is¡¡said£»
or¡¡a¡¡process¡¡of¡¡evolution£»¡¡must¡¡have¡¡occurred£»¡¡or¡¡else¡¡the¡¡whole
story¡¡must¡¡be¡¡given¡¡up£»¡¡as¡¡not¡¡only¡¡devoid¡¡of¡¡any¡¡circumstantial
evidence£»¡¡but¡¡contrary¡¡to¡¡such¡¡evidence¡¡as¡¡exists¡£
I¡¡placed¡¡before¡¡you¡¡in¡¡a¡¡few¡¡words£»¡¡some¡¡little¡¡time¡¡ago£»¡¡a
statement¡¡of¡¡the¡¡sum¡¡and¡¡substance¡¡of¡¡Milton's¡¡hypothesis¡£
Let¡¡me¡¡now¡¡try¡¡to¡¡state¡¡as¡¡briefly£»¡¡the¡¡effect¡¡of¡¡the
circumstantial¡¡evidence¡¡bearing¡¡upon¡¡the¡¡past¡¡history¡¡of¡¡the
earth¡¡which¡¡is¡¡furnished£»¡¡without¡¡the¡¡possibility¡¡of¡¡mistake£»
with¡¡no¡¡chance¡¡of¡¡error¡¡as¡¡to¡¡its¡¡chief¡¡features£»¡¡by¡¡the
stratified¡¡rocks¡£¡¡What¡¡we¡¡find¡¡is£»¡¡that¡¡the¡¡great¡¡series¡¡of
formations¡¡represents¡¡a¡¡period¡¡of¡¡time¡¡of¡¡which¡¡our¡¡human
chronologies¡¡hardly¡¡afford¡¡us¡¡a¡¡unit¡¡of¡¡measure¡£¡¡I¡¡will¡¡not
pretend¡¡to¡¡say¡¡how¡¡we¡¡ought¡¡to¡¡estimate¡¡this¡¡time£»¡¡in¡¡millions
or¡¡in¡¡billions¡¡of¡¡years¡£¡¡For¡¡my¡¡purpose£»¡¡the¡¡determination¡¡of
its¡¡absolute¡¡duration¡¡is¡¡wholly¡¡unessential¡£¡¡But¡¡that¡¡the¡¡time
was¡¡enormous¡¡there¡¡can¡¡be¡¡no¡¡question¡£
It¡¡results¡¡from¡¡the¡¡simplest¡¡methods¡¡of¡¡interpretation£»¡¡that
leaving¡¡out¡¡of¡¡view¡¡certain¡¡patches¡¡of¡¡metamorphosed¡¡rocks£»¡¡and
certain¡¡volcanic¡¡products£»¡¡all¡¡that¡¡is¡¡now¡¡dry¡¡land¡¡has¡¡once
been¡¡at¡¡the¡¡bottom¡¡of¡¡the¡¡waters¡£¡¡It¡¡is¡¡perfectly¡¡certain¡¡that£»
at¡¡a¡¡comparatively¡¡recent¡¡period¡¡of¡¡the¡¡world's¡¡historythe
Cretaceous¡¡epochnone¡¡of¡¡the¡¡great¡¡physical¡¡features¡¡which¡¡at
present¡¡mark¡¡the¡¡surface¡¡of¡¡the¡¡globe¡¡existed¡£¡¡It¡¡is¡¡certain
that¡¡the¡¡Rocky¡¡Mountains¡¡were¡¡not¡£¡¡It¡¡is¡¡certain¡¡that¡¡the
Himalaya¡¡Mountains¡¡were¡¡not¡£¡¡It¡¡is¡¡certain¡¡that¡¡the¡¡Alps¡¡and¡¡the
Pyrenees¡¡had¡¡no¡¡existence¡£¡¡The¡¡evidence¡¡is¡¡of¡¡the¡¡plainest
possible¡¡character¡¡and¡¡is¡¡simply¡¡this£ºWe¡¡find¡¡raised¡¡up¡¡on¡¡the
flanks¡¡of¡¡these¡¡mountains£»¡¡elevated¡¡by¡¡the¡¡forces¡¡of¡¡upheaval
which¡¡have¡¡given¡¡rise¡¡to¡¡them£»¡¡masses¡¡of¡¡Cretaceous¡¡rock¡¡which
formed¡¡the¡¡bottom¡¡of¡¡the¡¡sea¡¡before¡¡those¡¡mountains¡¡existed¡£
It¡¡is¡¡therefore¡¡clear¡¡that¡¡the¡¡elevatory¡¡forces¡¡which¡¡gave¡¡rise
to¡¡the¡¡mountains¡¡operated¡¡subsequently¡¡to¡¡the¡¡Cretaceous¡¡epoch£»
and¡¡that¡¡the¡¡mountains¡¡themselves¡¡are¡¡largely¡¡made¡¡up¡¡of¡¡the
materials¡¡deposited¡¡in¡¡the¡¡sea¡¡which¡¡once¡¡occupied¡¡their¡¡place¡£
As¡¡we¡¡go¡¡back¡¡in¡¡time£»¡¡we¡¡meet¡¡with¡¡constant¡¡alternations¡¡of¡¡sea
and¡¡land£»¡¡of¡¡estuary¡¡and¡¡open¡¡ocean£»¡¡and£»¡¡in¡¡correspondence¡¡with
these¡¡alternations£»¡¡we¡¡observe¡¡the¡¡changes¡¡in¡¡the¡¡fauna¡¡and
flora¡¡to¡¡which¡¡I¡¡have¡¡referred¡£
But¡¡the¡¡inspection¡¡of¡¡these¡¡changes¡¡gives¡¡us¡¡no¡¡right¡¡to¡¡believe
that¡¡there¡¡has¡¡been¡¡any¡¡discontinuity¡¡in¡¡natural¡¡processes¡£
There¡¡is¡¡no¡¡trace¡¡of¡¡general¡¡cataclysms£»¡¡of¡¡universal¡¡deluges£»
or¡¡sudden¡¡destructions¡¡of¡¡a¡¡whole¡¡fauna¡¡or¡¡flora¡£
The¡¡appearances¡¡which¡¡were¡¡formerly¡¡interpreted¡¡in¡¡that¡¡way¡¡have
all¡¡been¡¡shown¡¡to¡¡be¡¡delusive£»¡¡as¡¡our¡¡knowledge¡¡has¡¡increased
and¡¡as¡¡the¡¡blanks¡¡which¡¡formerly¡¡appeared¡¡to¡¡exist¡¡between¡¡the
different¡¡formations¡¡have¡¡been¡¡filled¡¡up¡£¡¡That¡¡there¡¡is¡¡no
absolute¡¡break¡¡between¡¡formation¡¡and¡¡formation£»¡¡that¡¡there¡¡has
been¡¡no¡¡sudden¡¡disappearance¡¡of¡¡all¡¡the¡¡forms¡¡of¡¡life¡¡and
replacement¡¡of¡¡them¡¡by¡¡others£»¡¡but¡¡that¡¡changes¡¡have¡¡gone¡¡on
slowly¡¡and¡¡gradually£»¡¡that¡¡one¡¡type¡¡has¡¡died¡¡out¡¡and¡¡another¡¡has
taken¡¡its¡¡place£»¡¡and¡¡that¡¡thus£»¡¡by¡¡insensible¡¡degrees£»¡¡one¡¡fauna
has¡¡been¡¡replaced¡¡by¡¡another£»¡¡are¡¡conclusions¡¡strengthened¡¡by
constantly¡¡increasing¡¡evidence¡£¡¡So¡¡that¡¡within¡¡the¡¡whole¡¡of¡¡the
immense¡¡period¡¡indicated¡¡by¡¡the¡¡fossiliferous¡¡stratified¡¡rocks£»
there¡¡is¡¡assuredly¡¡not¡¡the¡¡slightest¡¡proof¡¡of¡¡any¡¡break¡¡in¡¡the
uniformity¡¡of¡¡Nature's¡¡operations£»¡¡no¡¡indication¡¡that¡¡events
have¡¡followed¡¡other¡¡than¡¡a¡¡clear¡¡and¡¡orderly¡¡sequence¡£
That£»¡¡I¡¡say£»¡¡is¡¡the¡¡natural¡¡and¡¡obvious¡¡teaching¡¡of¡¡the
circumstantial¡¡evidence¡¡contained¡¡in¡¡the¡¡stratified¡¡rocks¡£¡¡I
leave¡¡you¡¡to¡¡consider¡¡how¡¡far£»¡¡by¡¡any¡¡ingenuity¡¡of
interpretation£»¡¡by¡¡any¡¡stretching¡¡of¡¡the¡¡meaning¡¡of¡¡language£»¡¡it
can¡¡be¡¡brought¡¡into¡¡harmony¡¡with¡¡the¡¡Miltonic¡¡hypothesis¡£
There¡¡remains¡¡the¡¡third¡¡hypothesis£»¡¡that¡¡of¡¡which¡¡I¡¡have¡¡spoken
as¡¡the¡¡hypothesis¡¡of¡¡evolution£»¡¡and¡¡I¡¡purpose¡¡that£»¡¡in¡¡lectures
to¡¡come£»¡¡we¡¡should¡¡discuss¡¡it¡¡as¡¡carefully¡¡as¡¡we¡¡have¡¡considered
the¡¡other¡¡two¡¡hypotheses¡£¡¡I¡¡need¡¡not¡¡say¡¡that¡¡it¡¡is¡¡quite
hopeless¡¡to¡¡look¡¡for¡¡testimonial¡¡evidence¡¡of¡¡evolution¡£¡¡The¡¡very
nature¡¡of¡¡the¡¡case¡¡precludes¡¡the¡¡possibility¡¡of¡¡such¡¡evidence£»
for¡¡the¡¡human¡¡race¡¡can¡¡no¡¡more¡¡be¡¡expected¡¡to¡¡testify¡¡to¡¡its¡¡own
origin£»¡¡than¡¡a¡¡child¡¡can¡¡be¡¡tendered¡¡as¡¡a¡¡witness¡¡of¡¡its¡¡own
birth¡£¡¡Our¡¡sole¡¡inquiry¡¡is£»¡¡what¡¡foundation¡¡circumstantial
evidence¡¡lends¡¡to¡¡the¡¡hypothesis£»¡¡or¡¡whether¡¡it¡¡lends¡¡none£»¡¡or
whether¡¡it¡¡controverts¡¡the¡¡hypothesis¡£¡¡I¡¡shall¡¡deal¡¡with¡¡the
matter¡¡entirely¡¡as¡¡a¡¡question¡¡of¡¡history¡£¡¡I¡¡shall¡¡not¡¡indulge¡¡in
the¡¡discussion¡¡of¡¡any¡¡speculative¡¡probabilities¡£¡¡I¡¡shall¡¡not
attempt¡¡to¡¡show¡¡that¡¡Nature¡¡is¡¡unintelligible¡¡unless¡¡we¡¡adopt
some¡¡such¡¡hypothesis¡£¡¡For¡¡anything¡¡I¡¡know¡¡about¡¡the¡¡matter£»¡¡it
may¡¡be¡¡the¡¡way¡¡of¡¡Nature¡¡to¡¡be¡¡unintelligible£»¡¡she¡¡is¡¡often
puzzling£»¡¡and¡¡I¡¡have¡¡no¡¡reason¡¡to¡¡suppose¡¡that¡¡she¡¡is¡¡bound¡¡to
fit¡¡herself¡¡to¡¡our¡¡notions¡£
I¡¡shall¡¡place¡¡before¡¡you¡¡three¡¡kinds¡¡of¡¡evidence¡¡entirely¡¡based
upon¡¡what¡¡is¡¡known¡¡of¡¡the¡¡forms¡¡of¡¡animal¡¡life¡¡which¡¡are
contained¡¡in¡¡the¡¡series¡¡of¡¡stratified¡¡rocks¡£¡¡I¡¡shall¡¡endeavour
to¡¡show¡¡you¡¡that¡¡there¡¡is¡¡one¡¡kind¡¡of¡¡evidence¡¡which¡¡is¡¡neutral£»
which¡¡neither¡¡helps¡¡evolution¡¡nor¡¡is¡¡inconsistent¡¡with¡¡it¡£
I¡¡shall¡¡then¡¡bring¡¡forward¡¡a¡¡second¡¡kind¡¡of¡¡evidence¡¡which
indicates¡¡a¡¡strong¡¡probability¡¡in¡¡favour¡¡of¡¡evolution£»¡¡but¡¡does
not¡¡prove¡¡it£»¡¡and£»¡¡lastly£»¡¡I¡¡shall¡¡adduce¡¡a¡¡third¡¡kind¡¡of
evidence¡¡which£»¡¡being¡¡as¡¡complete¡¡as¡¡any¡¡evidence¡¡which¡¡we¡¡can
hope¡¡to¡¡obtain¡¡upon¡¡such¡¡a¡¡subject£»¡¡and¡¡being¡¡wholly¡¡and
strikingly¡¡in¡¡favour¡¡of¡¡evolution£»¡¡may¡¡fairly¡¡be¡¡called
demonstrative¡¡evidence¡¡of¡¡its¡¡occurrence¡£
LECTURES¡¡ON¡¡EVOLUTION
II
THE¡¡HYPOTHESIS¡¡OF¡¡EVOLUTION¡£¡¡THE¡¡NEUTRAL¡¡AND
THE¡¡FAVOURABLE¡¡EVIDENCE¡£
In¡¡the¡¡preceding¡¡lecture¡¡I¡¡pointed¡¡out¡¡that¡¡there¡¡are¡¡three
hypotheses¡¡which¡¡may¡¡be¡¡entertained£»¡¡and¡¡which¡¡have¡¡been
entertained£»¡¡respecting¡¡the¡¡past¡¡history¡¡of¡¡life¡¡upon¡¡the¡¡globe¡£
According¡¡to¡¡the¡¡first¡¡of¡¡these¡¡hypotheses£»¡¡living¡¡beings£»¡¡such
as¡¡now¡¡exist£»¡¡have¡¡existed¡¡from¡¡all¡¡eternity¡¡upon¡¡this¡¡earth¡£
We¡¡tested¡¡that¡¡hypothesis¡¡by¡¡the¡¡circumstantial¡¡evidence£»¡¡as¡¡I
called¡¡it£»¡¡which¡¡is¡¡furnished¡¡by¡¡the¡¡fossil¡¡remains¡¡contained¡¡in
the¡¡earth's¡¡c
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